9 Genera (3 on Campus)
800 species
I. Leaves
A. Deciduous or (Persistent - some Quercus)
B. Alternate
C. Simple
D. Stipulate
E. Usually pinnate venation
II. Flowers
A. Monoecious
B. Axillary staminate aments or heads (Fagus)
C. Axillary pistillate spikes
1. Alone (Fagus, Quercus)
2. At base of staminate ament (Castanea)
II. Fruit
A. Nut with basal involucre
(cupule)
IV. Economic Importance
A. # 1 Hardwood Lumber (Floors, Furniture, Veneer)
B. Bark ->tannins
C. Edible fruits
D. Ornamental
E. Cork from Quercus suber
bark
V. Range
A. Fagus & Castanea = Eastern N. America
B. Quercus Most of USA & S. Canada
C. Lithocarpus & Castanopsis
Western N. America
Generic comparison
I. Fagus (Beech)
Latin for Beech
A. Triangular Nut enclosed
in friendly spiney involucre
II. Castanea (Chestnut/Chinkapin)
Latin for Chestnut
A. Round Nut(s) enclosed
in very unfriendly spiney involucre
III. Quercus (Oak)
Latin for Oak
A. Round/Ellipsoidal Nut enclosed entirely or partially in scaley
involucre
Species comparison
Fagus (Beech = “book” ;Gutenberg )
I. Shared characteristics
A. Smooth blue/gray bark (graffiti tree)
B. Acuminate apex; cuneate base; elliptical to oblong-ovate
D. Long spindle shaped bud on slender zigzag twig
E. Wood: red/brown; medium density; fine texture; uneven
figure
II. F grandifolia (American beech)
[probably because at maturity its crown is grandious]
it rots from the inside,
so don’t grow it next to your house
A. Serrate margins
B. > 10 pairs of lateral
veins
III. F. sylvatica (European beech)
[of the forest- watch out
for wild pigs]
A. Serrulate/Repand/Entire margins
B < 10 pairs of lateral veins
C. Leaves?, Buds? , Bark
is darker
Castanea (Chinkapin/Chestnut)
I. C. dentata (American Chestnut R.I.P)
[toothed]
A. Chestnut Blight (Cryphonectria parasitica) 1904
B. Light brown, light, course
texture, even figure wood
II. C. molissima (Chinese Chestnut)
A. Coarsely & sharply serrate leaf margins
B. Acuminate apex
C. Acute base
D. Nut enclosed in valved involucre covered with long very
sharp prickles
Quercus
Two Subgenera
I. Cyclobalanopsis (Cycle-cup oaks in Asia)
II. Quercus (Scale-cup oaks in Northern Hemisphere)
Three sections within Quercus (2 on campus)
I. Quercus = Leucobalanus (White Oaks)
A. No bristle tips on leaves
B. Glabrous inner surface of acorn shell
C. Sweet seeds
D. Whitish brown, dense, fine texture, uneven figure wood
E. Tyloses in vessels
II. Lobatae = Erythrobalanus (Red/Black Oaks)
A. Bristle tips on leaf lobes/teeth (mucronate)
B.Tomentose inner surface of acron shell
C. Bitter seeds
D. Reddish brown, dense, course texture, uneven figure wood
E. No tyloses in vessels
White Oaks
I. Lobed leaves
A. Glabrous
1. Q. alba (White oak)
a. cuneate leaf base
b. ovoid-oblong nut with warty 1/4 cupule
with short/sessile peduncle
c. light gray bark subdivided into rectangular plates
2. Q. robur (English oak)
a. auriculate leaf base
b. elongated oblong nut with scaley 1/3 cupule
with long slender peduncle
c. dark gray ridge/furrow bark
B. Pubescent
1. Q. macrocarpa (bur oak)
a. obovate leaf
b. big nut with 1/2 or more fringed cupule
c. dark gray vertical ridge/furrow bark
d. twigs often have corky ridges
2. Q. lyrata (overcup oak)
a. oblong-ovate leaf with wide middle lobes
[lyre-shaped]
b. cupule almost completely covers nut - no fringe
c. brownish gray rough irregular plate bark
II. Toothed leaves
A. 5-8 pairs of lateral veins
not all ending in teeth
1. Q. cerris (Turkey oak)
a. terminal & lateral buds with thread like stipules
b. elongated nut 1/2 fringed cupule sessile or with
short peduncle
2. Q. bicolor (Swamp white oak)
a. stipules not thread like
b. oblong-ovate nut 1/3 scale cupule on erect
2-6 cm peduncle, often paired
B. 9-13 pairs of lateral
veins all ending in teeth
1. Q. Muehlenbergii (Chinquapin oak)
a. sharp teeth curving toward apex
b. light gray, thin, fissured, scaly bark
2. Q. prinus (Chestnut oak) {Q. montana}
[Grk = oak]
a. blunt teeth, slightly or not curving toward apex
b. dark gray, thick, deeply furrowed into broad and
narrow ridges with orange inner bark visible in
fissures
Red Oaks
I. Entire Leaves
A. Q. imbricaria (Shingle oak)
[overlapping]
1. 2-5 cm wide
2. pubescent on underside
B. Q.
phellos (Willow oak)
[corky]
1. 0.5-2 cm wide
2. glabrous on underside
II. Lobed leaves
A. Conspicuous axillary tufts
of hairs
1. Q. palustris (Pin oak)
[swampy land]
a. reddish-brown twigs
b. reddish-brown buds
2. Q. Shumardii (Shumard oak)
a. gray -> grayish-brown twigs
b. gray ->grayish-brown buds
B. Inconspicuous axillary
tufts of hairs
1. Dull above, glaucous below, 7-11 lobes
a. Q. rubra (Northern Red oak)
1. shallow saucer shaped cupule
2. Shiny above, light yellowish-green below, 5-9 lobes
a. Q. velutina (Black oak)
[velvety]
1. yellowish orange inner bark
2. angled pentagonal 6-12 mm terminal buds
b. Q. coccinea (Scarlet oak)
1. reddish inner bark
2. rounded pentagonal 3-6 mm terminal buds
3. conspicuous concentric rings around
acorn apex